Aquarium

Angelfish (Scalar) Guide: Care, types, reproduction, and compatibility

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By Milthon Lujan

Angelfish (Pterophyllum scalare) at the Montreal Biodome. Source: mendel
Angelfish (Pterophyllum scalare) at the Montreal Biodome. Source: mendel

The angelfish, also popularly known as the scalar fish, is undoubtedly one of the jewels of freshwater aquariums. Its elegant silhouette, marked by long fins and a disc-shaped body, along with the incredible diversity of colors and patterns, has made it an undisputed favorite for aquarists of all levels.

Native to the Amazon basin in South America, these fish not only bring beauty but also fascinating behavior. However, for them to thrive and display their full splendor, it is crucial to understand their specific needs.

In this AquaHoy guide, we will take you step-by-step through everything you need to know. From how to set up the ideal aquarium and clarify doubts about its size, to a complete guide on fish compatibility, their diet, the spectacular varieties that exist, and the secrets to achieving successful breeding.

Angelfish taxonomy

  • Kingdom: Animalia
  • Phylum: Chordata
  • Class: Actinopterygii
  • Order: Cichliformes
  • Family: Cichlidae
  • Tribe: Heroini
  • Genus: Pterophyllum
  • Species: Pterophyllum altum (the largest), Pterophyllum leopoldi (the rarest), and Pterophyllum scalare (the most common)
  • Common Spanish names: Pez ángel, Pez escalar
  • English names: angelfish, angel fish.

Madireddy (2022) represented the first de novo sequencing and assembly of the complete genome of the freshwater angelfish (Pterophyllum scalare); the final genome assembly consisted of 15,486 contigs.

Characteristics and types of Angelfish

Angelfish are a cichlid species native to the Amazon River system, which is why they can be found in several South American countries such as Colombia, Guyana, French Guiana, Peru, and Brazil.

The unique appearance of the angelfish comes from its long fins that make it incomparable when compared to other ornamental fish.

On the other hand, it is difficult to differentiate male angelfish from females. The physical differences that can be used to distinguish the sexes are often only visible during spawning.

Angelfish are also characterized by their longevity; when compared to other ornamental fish, they can live for up to 12 years.

The three species of angelfish are P. scalare, P. altum, and P. leopoldi; however, the most common and commercially traded in aquarium stores is P. scalare.

Pterophyllum scalare

The common angelfish. It is the most widespread in the hobby, with countless varieties developed through selective breeding. The P. scalare angelfish can reach a body length of 15 cm and a height (including fins) of 20 cm.

Pterophyllum scalare - "Rio Nanay" - Aquarium tropicale du palais de la porte dorée Paris, France. Source: Cedricguppy - Loury Cédric
Pterophyllum scalare – “Rio Nanay” – Aquarium tropicale du palais de la porte dorée Paris, France. Source: Cedricguppy – Loury Cédric

Pterophyllum altum

The altum angelfish. It is larger, with a taller body and a distinctive notch above its snout. It is more demanding regarding water quality and is considered a challenge for advanced aquarists. P. altum can grow up to 18 cm in length and 25 to 33 cm in height.

Pair of angelfish Pterophyllum altum. Source: Jeff Kubina.
Pair of angelfish Pterophyllum altum. Source: Jeff Kubina.

Pterophyllum leopoldi

The smallest and most aggressive of the three, recognizable by a black spot at the base of the dorsal fin.

Popular angelfish varieties

Angelfish breeders are continuously developing new angelfish colors and patterns, but the best-known varieties include silver, veil, koi, zebra, marble, albino, leopard, and platinum. Likewise, a transgenic angelfish has also been developed. Below is a brief description of each variety:

  • Silver: The closest to the wild form, with vertical black bars.
  • Marble: Irregular patterns of black and white (or gold).
  • Black: An almost complete velvety black color.
  • Koi: Orange, black, and white spots, reminiscent of koi carp.
  • Zebra: Similar to silver but with more black stripes.
  • Albino: Without pigmentation, with red eyes.
  • Veil: With extremely long and flowing dorsal, anal, and caudal fins.
  • Ghost: A fish with almost no stripes, very pale.

Angelfish reproduction

Angelfish reproduction is a spectacle. They are devoted parents who care for their spawn.

Reproduction genetics

The results of the study by Chu et al. (2025) demonstrate that the dnd1 gene is essential for fertility and plays a crucial role in promoting the female sexual fate in the freshwater angelfish (Pterophyllum scalare).

Pair formation

The ideal is to let a pair form naturally from a group of juveniles. You will know you have a pair when they isolate themselves from the group and defend a territory. Differentiating males from females is difficult, but in adults, the male usually has a small nuchal hump on its head and its ovipositor tube is pointed, while the female’s is more rounded.

Depending on aquarium conditions, P. scalare reaches sexual maturity between six and twelve months. For angelfish reproduction, you should choose a male and a female and place them in their tank for spawning.

On the other hand, Luna-Figueroa and Arce (2023) highlight that providing live food before and during the reproductive stage results in more intense and brilliant coloration, better condition, immunological strength, more active swimming, more numerous and frequent spawns, and a larger number of fry.

Spawning

Usually, three to four hours before reproduction, the male and female scalar fish clean, using their mouths, the substrate where they will lay the eggs (Thilakarathne et al., 2021). The pair will clean a smooth surface (a broad leaf, a log, or a breeding cone) and the female will deposit rows of angelfish eggs, which the male will immediately fertilize.

The female angelfish can lay hundreds of eggs every one or two weeks if the eggs are removed from the tank. A successful spawn can produce up to 1000 eggs. The eggs are generally deposited on vertical surfaces such as a leaf, the filter tube, or a section of the aquarium wall. Sihombing and Sitanggang (2021) evaluated different types of substrate and concluded that the ceramic substrate is the best for the scalar fish to deposit their eggs.

Pair of angelfish guarding their eggs. Source: 5snake5
Pair of angelfish guarding their eggs. Source: 5snake5

Care of the spawn

Both parents will fan the eggs with their fins to oxygenate them and will remove any that are not viable (those that turn white). Depending on the tank’s water temperature, the eggs will hatch in two to three days.

Care of the fry

After hatching, the fry will remain attached to the surface, feeding on their yolk sac. After 5-7 days, they will begin to swim freely. This is the time to feed them with infusoria or newly hatched brine shrimp. Eiras et al. (2019) report that angelfish larvae can be raised in salinities of up to 4 g/L NaCl, and that the best growth occurs at 2 g/L NaCl, with a feeding frequency of 4 times per day.

Transporting juveniles

One of the most important activities is the transfer of angelfish from hatcheries to ornamental stores or the aquarist’s home; in this regard, you can use eugenol at a concentration of 15.9 mg/L to transport the fish for a period of 7 hours (de Oliveira et al., 2019).

For their part, de Oliveira et al. (2022) evaluated the effects of the essential oils of Lippia sidoides (EOLS) and Cymbopogon citratus (EOCC), concluding that the best concentrations for sedation and anesthesia for the angelfish (Pterophyllum scalare) for both essential oils were 10 and 25 mg L−1.

Feeding scalar fish

Freshwater angelfish are easy to feed and will take all types of fish food because they are omnivores. In nature, they feed on small crustaceans, insects, and plant matter. For vibrant health, their diet must be varied.

  • Base food: High-quality flakes or pellets for cichlids.
  • Live or frozen food: Essential for their development and to encourage reproduction. Options like artemia, mosquito larvae, daphnia, or tubifex are excellent. Luna-Figueroa and Arce (2023) cite that Artemia franciscana nauplii, Moina wierzejski, and Panagrellus redivivus showed the highest daily and specific growth rates in angelfish larvae and juveniles, respectively; however, Oktavia and Firmani (2024) found that the use of Daphnia sp. as an exclusive natural food is superior to artificial pellets and other combinations of natural food for the growth and survival of manfish (P. scalare).
  • Vegetable supplement: You can supplement with spirulina or homemade mashes that include peas or spinach.

The question “what do angelfish eat?” is answered with one word: variety. Feed them in small amounts twice a day.

Feed additives

The results of the research by Lima et al. (2024) demonstrated the potential of dietary supplementation with 0.2% β-glucans to improve the condition factor and positively modulate the intestinal microbiota of juvenile angelfish (P. scalare), without affecting other performance or blood parameters.

Feeding larvae and fry

For fry, Artemia nauplii are the best option to ensure rapid growth and higher survival. Likewise, Aziz et al. (2023) recommend feeding angelfish larvae with Artemia that has been enriched with microalgae (Tetraselmis sp. + Chlorella sp.); while Corecha et al. (2023) recommend the use of 75% Artemia franciscana nauplii and 25% Moina minuta neonates to reduce the feeding costs of angelfish larvae; and Eiras et al. (2022) report a feeding rate of 400 Moina sp. per postlarva/day, divided into two portions per day.

In the case of Pterophyllum scalare juveniles, Mao et al. (2023) recommend supplementing the commercial diet with 0.1% Artemia to improve the survival and growth of the fish, with a higher relative weight rate, standard length, and total length than the control group.

Feeding adult angelfish

In pet stores, you can find a variety of foods for your adult scalar fish; however, below we want to describe some research that can complement, and therefore improve, the nutrition of your angelfish.

Hamed et al. (2018) recommend the use of summer savory (Satureja hortensis) oil at an inclusion level of 200 mg/kg of feed to supplement the diet of angelfish, in order to improve growth, stress resistance, and the fish’s immune response.

Likewise, if you wish to enhance the coloration of your scalar fish, you can add 3% spirulina meal to the commercial feed, improving the color by a value of 3 (Lili et al., 2021).

Care of the Scalar fish

Angelfish can be kept in a wide variety of aquarium types, such as community and planted tanks; however, due to their height, angelfish need tall tanks. An adult angelfish requires space to swim vertically without its fins being damaged.

  • Absolute minimum for a pair: 120 liters, but a 200-liter aquarium is strongly recommended for a small group of 4-5 young specimens, allowing them to establish their hierarchy and form pairs naturally.
  • Height: The aquarium must be at least 50 cm tall.
  • General rule: Calculate about 40-50 liters of water for each adult angelfish.

In Table 01, we present the range that the main water quality parameters in your aquarium must meet in order to raise angelfish without any problems.

Table 01. Technical sheet for raising angelfish.

ParámetroRango
Volumen de la pecera200 litros para 4 adultos
Temperatura del agua25 a 28 oC.
pH6.0 – 8.0
Dureza del agua3 y 8o dKH (54 a 145 ppm)

Angelfish can be raised in a temperature range of 27 to 28 °C; however, Alvarez-Lee et al. (2023) suggest that an intermediate temperature of 28 °C should be used in production systems, as this condition represents the optimal physiology for the freshwater angelfish, and where the animals showed satisfactory performance.

Filtration and decoration

They prefer water with low current. An external or hang-on-back filter with regulated flow is ideal. Provide them with a fine and sandy substrate, and decorate with vertical driftwood and tall, broad-leaved plants like the Amazon Sword (Echinodorus), Vallisneria, or Anubias. These plants not only offer them security but are also the preferred place for them to deposit their eggs.

Angelfish behavior and compatibility

Generally, scalar fish are considered “semi-aggressive” because they chase each other in the aquarium. However, this territorial behavior occurs during the breeding season. dos Santos et al. (2018) conducted an experiment on the social interaction of angelfish and discovered that the increase in aggression is associated with the water change level; in this regard, the researchers recommend changing small volumes of water to prevent aggression and thus reduce the probability of injuries, stress, and affecting the fish’s well-being.

Environmental changes can affect some behavioral patterns in fish; in this sense, Ribeiro et al. (2021) evaluated the effects of water temperature on the aggressive behavior of male P. scalare, concluding that at low (23.8 °C) and high (31.7 °C) temperatures, there was a reduction in aggressive displays.

Fish compatible with scalars

The key is to choose tank mates of a similar size that are peaceful and not fin-nippers.

  • Medium-sized Tetras: Neon tetras (if introduced when the angelfish are young), cardinal tetras, bleeding heart tetras, or serpae tetras. They should be kept in schools of at least 8-10 individuals.
  • Corydoras and Ancistrus: Excellent bottom dwellers that will not compete for space. Keep them in groups of 6 or more.
  • Dwarf Cichlids: The Ramirezi (Mikrogeophagus ramirezi) or the Kribensis Cichlid can be good companions in large aquariums with enough space and hiding spots.
  • Loricariids: Moderately sized Plecos like Ancistrus are a great addition.

Warning! Species to avoid and risky combinations

  • Very small fish: Guppies, small neons, or galaxy rasboras can be devoured by adult angelfish.
  • Fin-nippers: Fish like the tiger barb or Buenos Aires tetra are a terrible choice, as they will harass the angelfish and damage their long fins.
  • Gouramis: Although some guides list them as compatible, we advise against this mix. Both can be territorial and have similar body shapes, which often leads to conflict.
  • Goldfish: They have completely different temperature and water requirements and should not be mixed.

Diseases of angelfish

Several common freshwater diseases affect angelfish in captivity:

Ich

White spot disease is caused by stress, which usually results from poor water conditions in the aquarium. The protozoan parasite Ichthyophthirius multifiliis causes white spots that attach to the fish’s body.

Ich can be treated by quarantining the affected fish in an aquarium where you should increase the water temperature to 30 °C with 1% salt (Abdullah-Al et al., 2021).

Dropsy

Dropsy is a symptom of an underlying bacterial infection, which can occur if the fish’s immune system is compromised.

This bacterial infection causes a bloated appearance, scales protruding from the body, bulging eyes, rapid breathing, lethargy, and loss of appetite.

You can treat dropsy with antibacterial medications recommended by a veterinarian, and add ⅛ of a teaspoon of salt for every five gallons of water.

Fin Rot

Fin rot is a bacterial infection that is commonly caused by fluctuating water parameters. This disease attacks the fins from the edges.

Several types of flavobacteria are responsible for fin rot, including Columnaris and Aeromonas.

You can treat fin rot by cleaning the aquarium and performing 20 to 50% water changes for at least two weeks.

Megalocytivirus

Johan et al. (2023) were the first to confirm the existence of different possible distribution routes of megalocytivirus in ornamental fish farms in Malaysia that raised angelfish (Pterophyllum scalare); they detected a high Megalocytivirus infection in P. scalare when water temperature, dissolved oxygen, and pH were elevated.

Parasites

Mora-Beltrán et al. (2024) reported the presence of metacercariae of the parasite Clinostomum sp. in a batch of Koi variety scalar fish (Pterophyllum scalare). They applied an antiparasitic treatment using a drug based on Praziquantel and Pyrantel Pamoate (Vermiplex® from ZOO Laboratories); this was administered in a combined manner: orally (25 mg total of Praziquantel and 75 mg total of Pyrantel Pamoate in a mash) and dissolved in the aquarium water (3 ml of dewormer dissolved in the water).

Conclusion

The angelfish is much more than just a pretty face in the aquarium. It is a species with complex and rewarding behavior that, with the right knowledge, can thrive and become the centerpiece of your home. By providing a suitable environment, a balanced diet, and compatible tank mates, you will enjoy the majesty of this incredible fish for many years to come.

Bibliographical References

ABDULLAH-AL MAMUN, M. D., NASREN, S., RATHORE, S. S., & RAHMAN, M. M. (2021). Mass infection of Ichthyophthirius multifiliis in two ornamental fish and their control measures. Annals of Biology, (2), 209-214.

Alvarez-Lee, L., Re, A. D., Diaz, F., Sanchez-Ovando, J. P., & Carrasco, L. P. (2023). Thermal biology and plasticity of the thermal metabolic scope of angelfish Pterophyllum scalare acclimated to different temperatures. International Aquatic Research, 15(3).

Aziz, N. S. A. N. A., Azani, N., Yatim, S. R., & Rasdi, N. W. (2023, March). Effects of mono and mix diets on growth of Artemia and its application as dietary sources of Pterophyllum scalare (Angelfish). In IOP Conference Series: Earth and Environmental Science (Vol. 1147, No. 1, p. 012011). IOP Publishing.

Corecha Fernandes Eiras Bruno José, Daniel Abreu Vasconcelos Campelo, Lorena Batista de Moura, Leonnan Carlos Carvalho de Oliveira, Liliane Marques de Sousa, Rauquírio Marinho da Costa, “Replacement of Artemia franciscana Nauplii by Moina minuta Neonates as Live Food on the Larviculture of Angelfish (Pterophyllum scalare -Schultze, 1823) and Severum (Heros severus – Heckel, 1840)“, Aquaculture Research, vol. 2023, Article ID 6118659, 8 pages, 2023. https://doi.org/10.1155/2023/6118659

Chu, W., Huang, S., Chang, C., Lin, Y., Wu, J., & Gong, H. (2025). Knockout of dead end 1 by CRISPR/Cas9 leads to loss of germ cells and male-biased sex development in freshwater angelfish (Pterophyllum scalare). Aquaculture, 599, 742180. https://doi.org/10.1016/j.aquaculture.2025.742180

de Oliveira, C. P. B., da Paixao Lemos, C. H., Felix, A., de Souza, S. A., Albinati, A. C. L., Lima, A. O., & Copatti, C. E. (2019). Use of eugenol for the anaesthesia and transportation of freshwater angelfish (Pterophyllum scalare). Aquaculture, 513, 734409.

de Oliveira, I.C., Oliveira, R.S.M., Lemos, C.H.d. et al. Essential oils from Cymbopogon citratus and Lippia sidoides in the anesthetic induction and transport of ornamental fish Pterophyllum scalare. Fish Physiol Biochem 48, 501–519 (2022). https://doi.org/10.1007/s10695-022-01075-3

dos Santos Gauy, A. C., Boscolo, C. N. P., & Gonçalves-de-Freitas, E. (2018). Less water renewal reduces effects on social aggression of the cichlid Pterophyllum scalare. Applied Animal Behaviour Science, 198, 121-126.

Eiras B. J. C. F., VerasG. C., AlvesA. X., & da CostaR. M. (2019). Effect of artificial seawater and feeding frequency on the larval culture of freshwater Amazonian ornamental fish banded cichlid Heros severus (Heckel, 1840) and angelfish Pterophyllum scalare (Schultze, 1823). Spanish Journal of Agricultural Research, 17(2), e0604. https://doi.org/10.5424/sjar/2019172-14645

Eiras, B. J. C. F., Campelo, D. A. V., de Moura, L. B., de Sousa, L. M., Nunes, I. S., de Oliveira, L. C. C., … & da Costa, R. M. (2022). Feeding rate and frequency during the first feeding of angelfish (Pterophyllum scalare-Schultze, 1823) and severum (Heros severus-Heckel, 1840) with Moina sp. Aquaculture, 553, 738106.

Hamed Ghafari Farsani, Mohammad Hasan Gerami, Mehdi Naderi Farsani, Ghasem Rashidiyan, Neda Mehdipour, Mona Ghanad & Caterina Faggio (2018): Effect of different levels of essential oils (Satureja hortensis) in diet on improvement growth, blood biochemical and immunity of Angelfish (Pterophyllum scalare Schultze, 1823), Natural Product Research, DOI: 10.1080/14786419.2018.1434635

Johan CAC, Abdullah MDD, Emilia SN, and Zainathan SC (2023) Molecular epidemiology of Megalocytivirus in freshwater angelfish (Pterophyllum scalare) from Johor, Malaysia, Veterinary World, 16(10):2158–2172.

Lili, W., Rizal, A., Herman, R. G., & Ramadhan, R. M. (2021). Effect of Spirulina flour on changes of color intensity in angelfish (Pterophyllum scalare) strain three color. Asian J Fish Aquat Res, 11(2), 52-58.

LIMA, E. C. S. D., SOUZA, F. P. D., FURLAN-MURARI, P. J., PANDOLFI, V. C. F., LEITE, N. G., MAINARDI, R. M., CHIDEROLI, R. T., PEREIRA, U. P., ARAÚJO, E. J. A., PUPIM, A. C. E., KOCH, J. F. A., & LOPERA-BARRERO, N. M.. (2024). Effects of dietary ß-glucans on the productive performance, blood parameters, and intestinal microbiota of angelfish (Pterophyllum scalare) juveniles. Anais Da Academia Brasileira De Ciências, 96(1), e20231006. https://doi.org/10.1590/0001-3765202420231006

Luna-Figueroa, J., & Arce, E. (2023). Alimentación del pez ángel Pterophyllum scalare con énfasis en el aprovechamiento de alimento vivo. Revista Iberoamericana de Ciencias. ISSN 2334-2501

Madireddy, I. (2022). First Ever Whole Genome Sequencing and De Novo Assembly of the Freshwater Angelfish, Pterophyllum scalare. Micropublication Biology, 2022.

Mao J, Chen Y, Lan S, et al. Growth and nutritional health of Pterophyllum scalare juveniles: Unleashing the benefits of feeding with Artemia sp. in aquariums. Israeli Journal of Aquaculture – Bamidgeh. 2023;75(2):1-13. doi:10.46989/001c.87423

Mora-Beltrán, N., Quintero-Pinto, L. G., Cruz Maldonado, O. A., & Cortés-Vecino, J. A. (2024). Reporte de caso: Infestación por Clinostomum sp. en Pterophyllum scalare y uso de Prazicuantel y Pamoato de Pirantel para su control. Revista Peruana De Investigación Agropecuaria, 3(1), e48. https://doi.org/10.56926/repia.v3i1.48

Oktavia, S. A. N., & Firmani, U. (2024). The Effect of Providing Different Natural Feeds on The Growth of Manfish (Pterophyllum scale). Jurnal Biologi Tropis, 24(1), 66–72. https://doi.org/10.29303/jbt.v24i1.6402

Rahmati-Holasoo H, Ebrahimzadeh Mousavi H, Nassiri A, Saeedi D, Ghojaye-Yousefi H, Taheri Mirghaed A. Investigating contamination with external and internal parasites in ornamental freshwater angelfish (Pterophyllum scalare). injvr 2024; 4 (2) :41-53 URL: http://injvr.com/article-1-68-en.html

Ribeiro, M. W. S., Oliveira, A. T., & Carvalho, T. B. (2021). Water temperature modulates social behavior of ornamental cichlid (Pterophyllum scalare) in an artificial system. Journal of Applied Aquaculture, 35(2), 410–422. https://doi.org/10.1080/10454438.2021.1973936

Sihombing, N. S., & Sitanggang, L. P. 2021. Effect of Use of Application Substrate Eggs Different Against Seed Productivity (Hatching Rate) Angelfish Fish (Pterophyllum Scalare). Berkala Perikanan Terubuk, 49(3), 1262-1271.

Thilakarathne, K. G. D. D., Hirimuthugoda, G. N., Lakkana, P. H. T., & Kumburegama, S. (2021). Embryonic and larval development in the freshwater angelfish (Pterophyllum scalare). Sri Lanka Journal of Aquatic Sciences, 26(1).

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